BlueGreen Labs was happy to co-author this work led by Alison Post. Alison showed that existing models of grassland phenology poorly predict green-up. Inclusion of precipitation, in addition to temperature, resulted in significantly better model results, although different thresholds apply across different climate regions. The models predict a green-up across most of the grassland sites studied. You can find the abstract and link to the full paper on the manuscript page.
Predicting spring green-up across diverse North American grasslands
Updating grassland GDD models using precipitation